Acoma
            Aegialia
            Aegidium
            Agenius
            Amphicoma
            Anaides
            Anomala
            Anthypna
            Antichira
            Aphodius
            Aphonus
            Aphotaenius
            Ataenius
            Ateuchus
            Calomacraspis
            Canthon
            Ceratophyus
            Cheirotonus
            Chlorata
            Chromovalgus
            Chrysina
            Chrysophora
            Clastocnemis
            Cloeotus
            Cnemida
            Coelodes
            Comythovalgus
            Copris
            Cratoscelis
            Cyclocephala
            Dasyvalgus
            Dialytes
            Diastictus
            Dicraeodon
            Didactylia
            Dilytellus
            Diplotaxis
            Dynastes
            Dyscinetus
            Enoplotrupes
            Euchirus
            Euparia
            Euparixia
            Euphoria
            Frickius
            Fruhstorferia
            Geotrupes
            Glaphyrus
            Glaresis
            Gnorimella
            Gnorimus
            Golinca
            Gymnetis
            Hapalonychus
            Heptaulacus
            Homonyx
            Homovalgus
            Hybalus
            Hybosorus
            Inca
            Incala
            Lachnosterna
            Lichnanthe
            Lichnia
            Liparochrus
            Lorditomaeus
            Macraspis
            Micraegialia
            Microvalgus
            Mimela
            Myodermum
            Ochodaeus
            Oncerus
            Onthophagus
            Orphnus
            Oryctomorphus
            Osmoderma
            Oxyomus
            Pachypus
            Pachystethus
            Paramacraspis
            Parastasia
            Paratrichius
            Peperonota
            Phaenomeris
            Phaeochrus
            Phanaeus
            Phyllophaga
            Platygenia
            Pleocoma
            Pleurophorus
            Plusiotina
            Plusiotis
            Polyphylla
            Popillia
            Proctophanes
            Propomacrus
            Psammobius
            Pseudomacraspis
            Ptenomela
            Rhyssemus
            Rutela
            Saprosites
            Serica
            Sisyphus
            Spilota
            Stegopterus
            Stenovalgus
            Strategus
            Strigoderma
            Synistovalgus
            Thyridium
            Toxocerus
            Trichiorhyssemus
            Trichiotinus
            Trichius
            Trigonopeltastes
            Typhoeus
            Valgus
Aphodiinae
Aphodiina
            Aphodius
Cetoniinae

            Aethiessa
            Agestrada
            Allorhina
            Amaurina
            Amithao
            Anatona
            Anatrinodia
            Anatropis
            Anochilia
            Anthracophora
            Aphanestes
            Argyripa
            Astraea
            Atrichelaphinis
            Atrichia
            Brachytrichia
            Bricoptis
            Calopotosia
            Campsiura
            Celidota
            Cetonia
            Cetonischema
            Chalcothea
            Charadronota
            Chiloloba
            Chlorobapta
            Chromoptilia
            Clinteria
            Clinteroides
            Clithria
            Coelodera
            Condradtia
            Conostethus
            Coptomia
            Cosmiomorpha
            Cotinis
            Cotinorrhina
            Cremastocheilus
            Cyclidinus
            Cyclidius
            Cymophorus
            Desicasta
            Diaphonia
            Dilochrosis
            Diplognatha
            Dirrhina
            Dischista
            Doryscelis
            Dysdiatheta
            Eccoptocnemis
            Epicometis
            Epixanthis
            Eriulus
            Erythroderma
            Eucetonia
            Euchilia
            Euchloropus
            Euchroea
            Euphoria
            Euphoriaspis
            Eupoecila
            Euryomia
            Euselates
            Gametis
            Genuchinis
            Genuchus
            Glycyphana
            Gnathocera
            Gnorimidia
            Gymentina
            Gymnetis
            Hemipharis
            Heterophana
            Heterosoma
            Hologymnetis
            Homothyrea
            Hoplopyga
            Hoplostomus
            Hypselogenia
            Ischipsopha
            Ischnostoma
            Iumnos
            Ixorida
            Leptothyrea
            Leucocelis
            Liocola
            Lissogenius
            Lomaptera
            Lomapterbides
            Lyraphora
            Macroma
            Macronota
            Macropodina
            Mausoleopsis
            Meroloba
            Metallesthes
            Micrelaphinis
            Micropeltus
            Micropoecila
            Mycterophallus
            Niphetophora
            Odontorrhina
            Oxycetonia
            Oxythyrea
            Pachnoda
            Paleopragma
            Pantolia
            Parachilia
            Phoxomela
            Poecilopharis
            Poecilophila
            Poecilophilides
            Polystigma
            Polystylactica
            Porphyronota
            Potosia
            Prionoryctes
            Progastor
            Protaetia
            Pseudinca
            Pseudochalcothea
            Pseudoprotaetia
            Pseudopygora
            Pygora
            Rhabdotis
            Rhinocoeta
            Rhomborrhina
            Rhyxiphloea
            Scaptobius
            Schizorrhina
            Sisyraphora
            Stenotarsia
            Stephanucha
            Stethodesma
            Systellorrhina
            Tapinoschema
            Temnorrhynchus
            Tephraea
            Tetragonorrhina
            Tetraodorrhina
            Thaumastopeus
            Tiarocera
            Tmesorrhina
            Torynorrhina
            Toxophyllus
            Trichaulax
            Trichostetha
            Trichothyrea
            Trigonophorus
            Trinodes
            Trinodia
            Tropinota
            Xenogenius
Golianthini
            Amaurodes
            Asthenorrhina
            Cheirolasia
            Chelorrhina
            Chondrorrhina
            Chordodera
            Coelorrhina
            Cyphonocephalus
            Diceros
            Dicranocephalus
            Dicronorrhina
            Dyspilophora
            Eudicella
            Genyodonta
            Goliathus
            Heterorrhina
            Mecynorrhina
            Megalorrhina
            Melinesthes
            Mycteristes
            Neptunides
            Pedinorrhina
            Plaesiorrhina
            Ptychodesthes
            Raceloma
            Ranzania
            Smaragdesthes
            Stephanorrhina
            Taeniesthes
            Theodosia
Coprinae

            Gromphas
Coprini
            Bdelyrus
            Canthidium
            Catharsius
            Chalcocopris
            Choeridium
            Copris
            Coptodactyla
            Coptorhina
            Dichotomius
            Heliocopris
            Holocephalus
            Ontherus
            Paraphytus
            Pedaria
            Pinotus
            Scatimus
            Synapsis
            Uroxys
Oniticellini
            Caccobius
            Cyptochirus
            Diastellopalpus
            Digitonthophagus
            Euoniticellus
            Helictopleurus
            Macropocopris
            Oniticellus
            Onthophagus
            Phalops
Onitini
            Bubas
            Chironitis
            Drepanocerus
            Onitis
Phanaeini
            Coprophanaeus
            Diabroctis
            Oxysternon
            Phanaeus
            Sulcophanaeus
Scarabaeini
            Anachalcos
            Ateuchus
            Canthochilum
            Canthon
            Canthonella
            Canthonosoma
            Cephalodesmius
            Circellium
            Deltochilum
            Epilissus
            Epirinus
            Eurysternus
            Gymnopleurus
            Mnematidium
            Mnematium
            Pachylomera
            Pachysoma
            Pseudocanthon
            Scarabaeus
            Sisyphus
            Temnoplectron
Dynastinae

            Adoryphorus
            Aegopsis
            Agaocephala
            Allomyrina
            Anastrategus
            Ancognatha
            Aneurystypus
            Anomolomorpha
            Anoplocephalus
            Aphonides
            Aphonus
            Archophileurus
            Augoderia
            Augosoma
            Bothynus
            Brachysiderus
            Chalcosoma
            Chalepidus
            Cheiroplatys
            Coelosis
            Corynophyllus
            Cryptodus
            Cyclocephala
            Cyphonistes
            Dasygnathus
            Dilobocerus
            Dipelicus
            Dynastes
            Dyscinetus
            Enema
            Eophileurus
            Epiphileurus
            Euetheola
            Eupatorus
            Golofa
            Haploscapanes
            Hatamus
            Hemiphileurus
            Heterogomphus
            Heteroligus
            Heteronychus
            Hexodon
            Homophileurus
            Hyphoryctes
            Isodon
            Ligyrus
            Liteupatorus
            Lonchotus
            Megaceras
            Megasoma
            Megasominus
            Metanastes
            Mimeoma
            Oryctes
            Oxygrylius
            Papuana
            Paraspidolea
            Pentodon
            Phileurus
            Phyllognathus
            Pimelopus
            Podischnus
            Pseudaphonus
            Pseudohomonyx
            Pseudopimelopus
            Pseudoryctes
            Ruteloryctes
            Scapanes
            Semanopterus
            Stenocrates
            Strategus
            Syrictes
            Temnorrhynchus
            Trichogomphus
            Trioplus
            Xyloryctes
            Xylotrupes
Geotrupinae
Geotrupini
            Athyreus
            Bolbelasmus
            Bolboceras
            Bolbocerosoma
            Eucanthus
            Kolbeus
            Odontaeus
Melolonthinae

            Ablabera
            Alobus
            Amphimallon
            Ancistrosoma
            Anisonyx
            Anomolochela
            Anoxia
            Antitrogus
            Apogonia
            Astaena
            Asthenopholis
            Athlia
            Autoserica
            Barybas
            Brachyllus
            Brahmina
            Calonota
            Castanochilus
            Ceraspis
            Chasme
            Chioneosoma
            Chlaenobia
            Chlorochiton
            Chnaunanthus
            Coenonycha
            Congella
            Coniopholis
            Cyclohirta
            Cyphochilus
            Dasyus
            Deuterocaulobius
            Diazus
            Dichelomorpha
            Dichelonyx
            Dichelus
            Dicrania
            Dinacoma
            Diphucephala
            Diplotaxis
            Ectinohoplia
            Elaphocera
            Empecta
            Enaria
            Encya
            Enthora
            Epholcis
            Eriesthis
            Eulepida
            Exopholis
            Frenchella
            Gama
            Geotrogus
            Gronocarus
            Gymnoloma
            Gymnopyge
            Haplidia
            Haplonycha
            Haplopsis
            Heptophylla
            Heterochelus
            Heteronyx
            Hilarianus
            Hilyotrogus
            Holotrichia
            Hoplia
            Hoplocnema
            Hoplosternus
            Hypopholis
            Hyposerica
            Isonychus
            Lachnosterna
            Lasioserica
            Lepidiota
            Lepisia
            Lepitrix
            Leucopholis
            Liogenys
            Liparetrus
            Listrochelus
            Macrodactylus
            Maechidius
            Maladera
            Manopus
            Megistophylla
            Melolontha
            Microplidus
            Microserica
            Microtrichia
            Miridiba
            Modialis
            Monochelus
            Neoserica
            Neso
            Ochrosidia
            Odontria
            Omocrates
            Pachycnema
            Paraserica
            Pegylis
            Peritrichia
            Philochlaenia
            Phobetus
            Phyllophaga
            Phyllotocus
            Phytalus
            Phytolaema
            Platychelus
            Plectris
            Plectrodes
            Podolasia
            Polyphylla
            Proagosternus
            Pseudacratus
            Pseudoholophylla
            Pseudotrematodes
            Psilopholis
            Rhinaspis
            Rhizotrogus
            Schizonycha
            Schonherria
            Serica
            Seriesthis
            Sparrmannia
            Stepanopholis
            Sulcipalpus
            Tanyproctus
            Thyce
            Tricholepis
            Xylonychus
Rutelinae
            Adoretosoma
            Adoretus
            Adorodocia
            Amblyterus
            Anisoplia
            Anomala
            Anomalepta
            Anoplognathus
            Antichira
            Areoda
            Bolax
            Brachysternus
            Callirhinus
            Callistethus
            Calloodes
            Chasmatopterus
            Chasmodia
            Cotalpa
            Cyclocephala
            Dyscinetus
            Epectinaspis
            Epichrysus
            Evanos
            Geniates
            Heterosternus
            Homaloplia
            Hoplognathus
            Hylamorpha
            Ischnopopillia
            Lagochile
            Leucothyreus
            Macraspis
            Macropoides
            Macropopillia
            Mimadoretus
            Mimela
            Oncerus
            Pachystethus
            Paracotalpa
            Parareoda
            Paraschizognathus
            Pelidnota
            Phyllopertha
            Platycoelia
            Plusiotis
            Pocalta
            Polymoechus
            Popillia
            Pseudadoretus
            Pseudoschizognathu
            Pseudosinghala
            Repsimus
            Rhombonalia
            Saulostomus
            Schizognathus
            Spilopopillia
            Spodochlamys
            Strigoderma
            Strigodermella
            Trigonochilus
            Triodonta
            Trochalus